Australia won the first match of a four-Test series by 118 runs on Monday, but the tourists were criticised for their ebullient celebration of the dramatic run out of De Villiers in the second innings.
De Villiers was run out for nought after being involved in a mix-up with opening batsman Aiden Markram, with Lyon, who had knocked off the bails, dropping the ball towards the sprawled batsman as he ran over to join ecstatic celebrations.
Lyon, who returned match figures of 3-136, admitted the offence and sanction proposed by match referee Jeff Crowe that also saw the 30-year-old penalised with a demerit point.
The bowler was said to have apologised to De Villiers and the pair were seen shaking hands while the teams warmed up today.
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