Then conservationists hoisted the limp bodies into crates, grasping big paws, thick tails and even the imposing heads of animals that earlier pounced on an antelope carcass used as bait to bring them closer to a veterinarian with a tranquiliser gun.
The operation this week to move seven South African lions to Rwanda in east-central Africa is an upbeat tale in an otherwise challenging picture for wildlife on the continent, where poaching has surged and human encroachment is shrinking habitats. Lions are designated as vulnerable on an international "red list" of species facing threats.
An Associated Press writer and photographer watched as crane operators loaded the lion crates onto a truck that arrived with two male lions from nearby Tembe Elephant Park. The vehicle then drove to Johannesburg for a charter flight to Rwanda.
The lions' new home is Akagera National Park, where cattle herders poisoned Rwanda's last lions about 15 years ago after the park was left unmanaged in the genocide's wake. Returning refugees took over much of the park, reducing its size by more than half to 1,122 square kilometers in 1997, according to African Parks, a Johannesburg-based group that manages Akagera and other national wildlife parks in Africa.
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At Phinda, the five lionesses bound for Rwanda were unexpectedly joined in their holding pen by three other lions that burrowed their way inside. Toft had to dart all eight before workers could enter to fasten satellite tracking collars on the five lions earmarked for travel to Rwanda.
The sedated lions looked vulnerable, jarring with the popular notion of lions as animal royalty. They were blindfolded with cloth and park staff and others moved quietly and spoke softly to reduce the chance of any lion awakening.